It is fairly certain that Ghana has been occupied by Negroid peoples since prehistoric times. Members of the Akan family, who make up some 44% of the population, include the Twi, or Ashanti, inhabiting the Ashanti Region and central Ghana, and the Fanti, inhabiting the coastal areas. In the southwest, the Nzima, Ahanta, Evalue, and other tribes speak languages related to Twi and Fanti. The Moshi-Dagomba constitute 16% of the population, the Ewe 13%, and the Ga 8%. The Accra plains are inhabited by tribes speaking variants of Ga, while east of the Volta River are the Ewe living in what used to be British-mandated Togoland. All these tribes are fairly recent arrivals in Ghana, the Akan having come between the 12th and 15th centuries, the Ga-Adangbe in the 16th century, and the Ewe in the 17th century. Most of the inhabitants of the Northern Region belong to the Mole-Dagbani group of Voltaic peoples or to the Gonja, who appear to bear some relation to the Akan. European and other groups account for only 0.2% of the population.