In 1996, Ecuador had 20 agricultural, medical, scientific, and technical learned societies and research institutes, most notably the General Directorate of Hydrocarbons, the Institute of Nuclear Sciences, the Ecuadoran Institute of Natural Sciences, and the National Institute of Agricultural Research, all in Quito; the Charles Darwin Research Station in the Galapagos Islands; and the Naval Oceanographic Institute In Guayaquil. Ecuador has 19 colleges and universities offering degrees in basic and applied sciences. In 1987–97, science and engineering students accounted for 27% of college and university enrollments. In 1987–97, Ecuador had 146 scientists and engineers and 42 technicians per million people engaged in research and development.