Colombia has an abundance of fish in its Caribbean and Pacific coastal waters and in its innumerable rivers. Lake Tota in Boyacá and Lake La Cocha in Nariño abound in trout, as do the artificial reservoirs of Neusa and Sisga in Cundinamarca. About half of the annual catch consists of freshwater fish. Tarpon are caught in the delta waters of the Magdalena, and sailfish, broadbills, and tuna in the Caribbean. The 2000 fish catch was 129,644 tons. The development of crustacean aquaculture has expanded since the mid-1980s, especially for Penaeid shrimps. Aquaculture production consists primarily of tilapia and rainbow trout.